1994	B.S., Department of Zoology, National Taiwan University
1996	M.S., Molecular Medicine Institute, College of Medicine, National Taiwan University, Taiwan.
2003	Ph.D., Department of Biology, New York University
6/03- 10/03	Postdoctoral Fellow, Department of Molecular and Cellular Biology, Breast Center, Baylor College of Medicine, USA.
11/03- 8/10	Postdoctoral Fellow, Department of Genetics, Harvard Medical School, USA.
09/10~present	Research Fellow, Institute of Molecular Biology, Academia Sinica